距今50多年前，一艘来自赛博坦的飞船坠落月球，由此引发了美苏两国的太空竞赛。时间回到21世纪初，汽车人终于挫败霸天虎的入侵，继而与人类合作，共同保卫美丽的地球。为了防止霸天虎找到能量柱为非作歹，擎天柱与战友飞赴月球，更从当年的飞船中救出了汽车人的先代领导者——御天敌。然而御天敌却和霸天虎有更大的阴谋和危机隐藏在那月影背后！
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娜奥米·斯科特,凯尔·加尔纳,德鲁·巴里摩尔,罗丝玛丽·德薇特,卢卡斯·盖奇,迪兰·格鲁拉,彼得·雅各布森,雷·尼科尔森,劳尔·卡斯提洛,达芙妮·泽利,迈尔斯·古铁雷斯-赖利,Zebedee Row,Delphi Harrington,Micaela Lamas,David Peter White
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